FRANKFURT, Germany — While Bayern Munich and Borussia Dortmund made no last-minute moves, Bundesliga rival Schalke and other clubs signed new players as the transfer deadline expired Wednesday with record spending and Hamburger SV landing Brazilian Olympic gold medalist Douglas Santos.

The five-year deal was signed shortly before the German transfer deadline ran out at 1600 GMT. Hamburg gave no details about the deal, but local media reports put the transfer at between 7 million and 10 million euros ($7.79 million and $11.13 million).

Douglas Santos, a left back, last played for Atletico Mineiro and was in Brazil’s lineup that beat Germany in the Olympic final.

Schalke’s latest addition to an already heavily revamped squad was attacking midfielder Yevhen Konoplyanka from Sevilla. The Ukraine international, who helped Sevilla win the Europa League last season, arrives on a one-season loan with an option to make the move more permanent.

“We are very pleased to have gotten a quick player who is a scoring threat and can play on both wings,” Schalke coach Markus Weinzierl said.

According to the German news agency dpa’s calculation, based on estimates, Bundesliga clubs spent 500 million euros ($556 million) for the first time by spending 506.5 million euros ($565.66 million) on new players. At the same time, the 18 clubs earned about 450 million euros ($502.56 million) from transfers, another all-time high.

Schalke reshuffled its squad following the departure of winger Leroy Sane for Manchester City, bringing in 16 new players while letting 10 go. Weinzierl is also a new arrival.

Konoplyanka reunites with former Sevilla defender Coke. But the right back had a serious right-knee injury on arrival and is not available.

Dortmund already had signed eight new players and made no new moves as the deadline approached.

Here’s a look at other major moves by Bundesliga clubs on the last day of the transfer window:

BAYER LEVERKUSEN

One club that gave up players was Leverkusen, a Champions League participant considered one of the contenders for a high finish in the Bundesliga. Leverkusen sent two defenders on loan to Bundesliga rivals. Kyriakos Papadopoulos went to RB Leipzig and Andre Ramalho moved to Mainz. Both clubs have an option to buy out their contracts.

Leverkusen signed Austria defender Aleksandar Dragovic ahead of the season.

WERDER BREMEN

After losing the Bundesliga opener to Bayern Munich 6-0, Bremen acquired midfielder Serge Gnabry from Arsenal. The Germany under-21 player was the top scorer for the silver medal-winning team at the Rio de Janeiro Olympics earlier this month.

At the same time, Bremen sent defender Leon Guwara on loan to Darmstadt.

RB LEIPZIG

The promoted club has sent defender Atinc Nukan back to Besiktas on loan. In addition to Papadopoulos, Leipzig has signed Brazilian defender Bernardo and Scottish midfielder Oliver Burke in recent days.

INGOLSTADT

Central defender Marcel Tisserand has arrived from Monaco on a four-year deal.

EINTRACHT FRANKFURT

Frankfurt has taken forward Shani Tarashaj from Everton on loan for one season. The Switzerland international is expected to take the place of Ante Rebic, who is out until further notice with mononucleosis.

WOLFSBURG

Wolfsburg added defender Philipp Wollscheid from Stoke on loan for this season, with an option to make it long-term. Wollscheid played for Nuremberg while Dieter Hecking, now the coach at Wolfsburg, was in charge there.

Wolfsburg said efforts to sign forward Son Heung-min from Tottenham had fallen through.